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TV020
Vegetation Data from inside and outside Elk Exclosures in the South Fork Hoh Permanent Reference Stands, Olympic National Forest , 1979-2007

CREATOR(S): Jerry F. Franklin, Mark E. Harmon
PRINCIPAL INVESTIGATOR(S): Mark E. Harmon
ORIGINATOR(S): Jerry F. Franklin
DATA SET CONTACT PERSON: Mark E. Harmon
METADATA CONTACT: Mark E. Harmon
ABSTRACTOR: Mark E. Harmon
METHOD CONTACT: Mark E. Harmon
FORMER INVESTIGATOR: Sarah E. Greene, Diane L. Mitchell, Miles Henstrom
DATA SET CREDIT:
This study was funded by the National Park Service, Andrews LTER, and Kaye and Ward Richardson endowment.
METADATA CREATION DATE:
5 Feb 2016
MOST RECENT METADATA REVIEW DATE:
5 Feb 2016
KEYWORDS:
community structure, plant cover, herbivory
PURPOSE:
This study was established following the 1978 pulse that was conducted on the South Fork of the Hoh River, Olympic National Park, Washington. During the pulse it became apparent that Roosevelt elk were having a major impact on understory plants and tree regeneration. Exclosures were established to exclude the elk. Comparisons of plots inside and outside the exclosures were to indicate the effect of the elk.
METHODS:
Experimental Design - TV020:
Description: Exclosures comprised of livestock fencing were established in two existing reference stands. The exclosures were 100 m by 50 m in size, with a height of approximately 3 m. A comparable area adjacent to each exclosure was also established as a control. In each area four 50 m transects were established: two end on end running along the central 100 m axis and two crossing the long axis 25 m from the start and end of 100 m length. Vegetation response in terms of herbs, shrubs, and tree regeneration was periodically measured inside and outside the exclosures. Initially the start and stop point of the transects was not marked. At the third measurement in 1981, wooden stakes were placed at the start and end of each transect. In 2007 these were replaced by PVC piping.
Field Methods - TV020:
Description:

Herb cover measurements. Cover of herbs, including mosses, liverworts, ferns, grasses, sedges, and other vascular plants were quantified using Daubenmire microplots (50 cm 20 cm) systematically spaced along the 50 m transect. Initially a microplot was placed at the start of each meter along the transect. After the first two samplings (spring and summer), the microplots were placed at the start of each odd meter (e.g., 1, 3, 5 m). It is likely that the microplots were consistently placed within a 50 cm in part due to variations in where the tape measure was initially placed, large unavoidable objects (e.g., logs, tip-up mounds), etc. Comparison of different measurements tend to show consistency in patches of the various species.

Cover was estimated using six classes >0 to 5, 5-25, 25-50, 50-75, 75-95, and >95%. Each species was estimated on its own, therefore total cover can exceed 100%. Mosses and liverworts were combined in some measurements, but in others the individual species were recorded to the extent possible. In some years, the cover of logs, bare soil, and other surface features were recorded in part to document disturbances to the ground surfaces.

Shrub cover. The start and stop point of shrubs was noted along the transect. In theory any shrub canopy crossing the 50 m transect was to be recorded. However, in a practical sense the cover was generally >10 cm in length. The way that vine maple was considered varied. In some cases all vine maple canopy was used including that of small trees far above the transect. In other cases, only the vine maple below head height was included. All other shrub species were consistently measured. There were difficulties in distinguishing several Vaccinium species (alaskaense and ovatum). These were lumped together as Vaccinium alaskaense).

Tree regeneration. Tree seedlings and saplings less than 5 cm diameter at breast height were counted along a 50 m by 1 m belt transect. The count was divided into six height classes: 10-25 cm, 25-50 cm, 50-75 cm, 75-100 cm, 100-150 cm, and >150 cm but less than 5 cm DBH. The count numbers are highly variable, in part because where the transect crossed logs (the primary tree regeneration location) varied and this could greatly influence the count. Attempts to mark the seedlings were not successful. The most variable counts were for the shortest size classes. These data should be used not so much to determine seedling, sapling density, as much as the relative proportions of species and size classes.

Models/Algorithms - TV020:
Description: To estimate the herb cover for a species on a transect (or part of transect), the mid-point of each cover class was used weighted by the number of microplots with that cover class. To estimate shrub cover, the start points were subtracted from the end points; these values were summed and divided by 50 m to get the proportional cover. The percent cover was proportional cover multiplied by 100. A list of species was compiled based on earlier measurement to help crews consistently identify and code species. Data were reorganized to run on various programs and in some cases were grouped into different starting community types representing different microsites (open and hydric, closed and mesic, open and mesic).
SITE DESCRIPTION:
South Fork of the Hoh River, Olympic National Park, Washington. The study sites are dominated by Sitka spruce-western hemlock forests growing on elevated river terraces. The climate is warm and wet, with up to 300 cm of annual precipitation.
TAXONOMIC SYSTEM:
None
GEOGRAPHIC EXTENT:
South Fork of the Hoh River, Olympic National Park, Washington. Covers an area of approximately 4 ha.
ELEVATION_MINIMUM (meters):
245
ELEVATION_MAXIMUM (meters):
250
MEASUREMENT FREQUENCY:
about 10 year intervals
PROGRESS DESCRIPTION:
Complete
UPDATE FREQUENCY DESCRIPTION:
irregular
CURRENTNESS REFERENCE:
Ground condition